German to English Other Livestock / Animal Husbandry Hunting

This is from an article about how introducing hunting could help improve animal numbers in African reserves.

The whole sentence is as follows:

Die Trophäenjagd ist besonders erfolgreich. Sie bringt den höchsten Ertrag bei gleichzeitig niedrigsten Entnahmeraten und – im Vergleich zu anderen Formen des Tourismus – geringen Investitionen und niedrigem Landverbrauch.

I can see two possible meanings, and I am not sure which is correct. My first thought was that, to go with high incomes, this must mean low outgoing costs....

Bearing in mind the topic of the article, I then thought that it could also mean something along the lines that the amount of animal wastage is very low. I.e.- other things mentioned in the article include raising animals for meat, for skins, for horn and bone....and in comparison to this, the number of animals killed by hunting would be relatively small.

Any ideas which is the correct meaning?