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Cartilage development
Further condensation involves change in matrix composition rich in type II collagen, type IX and XI collagen, Gla protein, aggrecan and link protein and downregulation of type I collagen. (...) Articular cartilage is rich in proteoglycans such as aggrecan (90% of the total proteoglycan content), link protein, decorin, cartilage derived oligomeric matrix protein, fibromodulin, lumican, biglycan, and perlecan. (...) Three SOX transcription factors (SOX9, SOX5 and SOX6) regulate early chondrogenic differentiation and expression of the ECM molecules, collagen (Col2a1, Col9a1), aggrecan, and link protein.