English to Spanish Science Zoology etología

¿Alguien puede decirme cómo se traducirían “ultimate” y “proximate” en un texto sobre biología del comportamiento (etología)? Os pongo ‘bastante’ contexto…

“Evolutionary questions are concerned with understanding behaviour in terms of its evolutionary history and function. Explanations of behaviour in this domain are called ***‘ultimate’***. ***Ultimate*** questions help us understand the adaptive value of behaviour (e.g., how does alarm calling increase the probability of survival?) and allow us to make predictions about change (e.g., in what ways should animals alter their foraging behaviour when a predator is present?). ***Ultimate*** causes are linked to evolutionary fitness, which in turn is closely related to individual reproductive output. Because population growth is a result of the cumulative reproduction of individuals, an understanding of ***ultimate*** causes may allow us to predict population-level consequences of specific changes in the environment

Mechanistic questions about behaviour deal with the ontogeny and control of behavioural patterns. Questions such as how does foraging behaviour develop in a young animal? and what is the neuronal control of cricket song? belong to this approach. These are also called ***‘proximate’*** questions19. In the conservation context, a detailed understanding of ***proximate*** mechanisms allows us to influence the behaviour of individuals and anticipate problems when animals are placed in novel situations (such as in captivity).
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***Ultimate*** and ***proximate*** considerations of the causes and consequences of mate choice (and sexual selection in general) yield new predictions and new opportunities in our efforts to preserve species and the habitats they live in."

GRACIAS de antemano.